Did human evolution favor individualists or altruists?

Eric Michael Johnson, writing for Slate:

Ultimately, [Ayn] Rand was searching for the origin of John Galt in the pages of human nature. But was she right? Are we rational egotists trapped in a net of social obligations? Or are we an innately social species for whom altruism was integral to our success on this planet? There was only one place she could look: the Pleistocene.

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